We introduce a class of uniformly $2$-nondegenerate CR hypersurfaces in $\mathbb{C}^N$, for $N>3$, having a rank $1$ Levi kernel. The class includes uncountably many examples of non-equivalent structures having a given constant nilpotent CR symbol for every such symbol. Using methods that are able to handle all cases with $N>5$ simultaneously, we solve the equivalence problem for the considered structures whose symbol is represented by a single Jordan block, classify their algebras of infinitesimal symmetries, and classify the locally homogeneous structures among them. We show that the remaining considered structures, which have symbols represented by a direct sum of Jordan blocks, can be constructed from the single block structures through simple linking and extension processes.