E4218 E4209 Proceedings of the National Academy of Sciences USA, 115, 18, pp. E4209-E4218 Proceedings of the National Academy of Sciences of the United States of America, 115(18), E4209-E4218. National Academy of Sciences Proceedings of the National Academy of Sciences USA, 115, E4209-E4218 Proceedings of the National Academy of Sciences of the United States of America, 115(18), E4209-E4218 Proceedings of the National Academy of Sciences of the United States of America 115 (2018) 18