Proceedings of the National Academy of Sciences of the United States of America Proceedings of the National Academy of Sciences of the United States of America, National Academy of Sciences, 2019, Epub ahead of print. ⟨10.1073/pnas.1810932116⟩ Proceedings of the National Academy of Sciences of the United States of America, 2019, Epub ahead of print. ⟨10.1073/pnas.1810932116⟩