Canadian Journal of Fisheries and Aquatic Sciences Canadian Journal of Fisheries and Aquatic Sciences, NRC Research Press, 2020, 77 (1), pp.164-176. ⟨10.1139/cjfas-2018-0444⟩ Canadian Journal of Fisheries and Aquatic Sciences, NRC Research Press, In press, ⟨10.1139/cjfas-2018-0444⟩