Tribe Aphodiini Leach, 1815 Type genus. Aphodius Hellwig, 1798 (see Alonso-Zarazaga & Krell 2011). See Dellacasa et al. (2001), and Smith & Skelley (2007) for synonymies and detailed description. The new taxon shares following synapomorphies with the members of Aphodiini: 1) labrum and mandibles invisible in antero-dorsal view; 2) scutellum well visible in dorsal view; 3) pronotum without any longitudinal groove; head without any longitudinal groove or costa; 4) protibia not perpendicularly truncate apically; 5) metatibial spurs separated by basal tarsomere; abdominal ventrites not fused, freely moveable.
Published as part of Kakizoe, Showtaro, Jiang, Ri-Xin, Cai, Hao-Hong & Wang, Shuo, 2021, Sphaeraphodius gaohani, a new genus and species of Aphodiini (Coleoptera Scarabaeidae: Aphodiinae) from Guangxi, China, pp. 333-340 in Zootaxa 4949 (2) on page 334, DOI: 10.11646/zootaxa.4949.2.6, http://zenodo.org/record/4636467
{"references":["Leach, W. E. (1815) Entomology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia. Vol. 9. Part 1. Blackwood, Edinburgh, pp. 57 - 172.","Alonso-Zarazaga, M. A. & Krell, F. T. (2011) Change of authorship of Aphodius and Oryctes to Hellwig, 1798 (Insecta: Coleoptera: Scarabaeidae). Zootaxa, 3060 (1), 67 - 68. https: // doi. org / 10.11646 / zootaxa. 3060.1.5","Dellacasa, G., Bordat, P. & Dellacasa, M. (2001) A revisional essay of world genus-group taxa of Aphodiinae. Memorie della Societa Entomologica Italiana, 79, 1 - 482.","Smith, A. B. & Skelley, P. E. (2007) A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa, 1458 (1), 1 - 80. https: // doi. org / 10.11646 / zootaxa. 1458.1.1"]}